Biosciences

College of Life and Environmental Sciences

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© The Author(s) 2018. Published by Oxford University Press. 1
This is an Open Access article distributed under the terms of the Creative Commons Attribution License (http://creativecommons.org/licenses/by/4.0/), which permits
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Except where otherwise noted, this item's licence is described as © The Author(s) 2018. Published by Oxford University Press. 1 This is an Open Access article distributed under the terms of the Creative Commons Attribution License (http://creativecommons.org/licenses/by/4.0/), which permits unrestricted reuse, distribution, and reproduction in any medium, provided the original work is properly cited