College of Life and Environmental Sciences
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Except where otherwise noted, this item's licence is described as © 2017 The Authors. Functional Ecology published by John Wiley & Sons Ltd on behalf of British Ecological Society. This is an open access article under the terms of the Creative Commons Attribution License (https://creativecommons.org/licenses/by/4.0/), which permits use, distribution and reproduction in any medium, provided the original work is properly cited.